Meet Gramma: The 141-year-old Galápagos tortoise died who witnessed two world wars died at the San Diego Zoo

San Diego Zoo mourns the loss of Gramma, a Galápagos tortoise estimated to be 141 years old. Born in the Galápagos Islands, she became a beloved "Queen of the Zoo" after arriving in the late 1920s. Her remarkable lifespan highlights the longevity of her species and the zoo's conservation efforts.

Meet Gramma: The 141-year-old Galápagos tortoise died who witnessed two world wars died at the San Diego Zoo
San Diego Zoo mourns the loss of Gramma, a Galápagos tortoise estimated to be 141 years old. Born in the Galápagos Islands, she became a beloved "Queen of the Zoo" after arriving in the late 1920s. Her remarkable lifespan highlights the longevity of her species and the zoo's conservation efforts.